Fruit bats in the genus Pteropus include the world's largest bat - the greater flying fox - which can have a wingspan of 180cm. They cannot echolocate, but don't really need to as their food (fruit and nectar) doesn't try to escape and because they roost in trees not in the pitch black of caves. Their eyesight alone is perfectly good enough for navigation. Fruit bats also have a very good sense of smell for detecting flowers and fruit.
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